taijasat tu vikurvanad
buddhih pranas ca taijasau
Translation by His Divine Grace A. C. Bhaktivedanta Swami Srila Prabhupada:
By further transformation of the mode of passion, the sense organs like the ear, skin, nose, eyes, tongue, mouth, hands, genitals, legs, and the outlet for evacuating, together with intelligence and living energy, are all generated.
Purport by His Divine Grace A. C. Bhaktivedanta Swami Srila Prabhupada:
The living condition in material existence depends more or less on one’s intelligence and powerful living energy. Intelligence to counteract the hard struggle for existence is assisted by the senses for acquiring knowledge, and the living energy maintains himself by manipulating the active organs, like the hands and legs. But on the whole, the struggle for existence is an exertion of the mode of passion. Therefore all the sense organs, headed by intelligence and the living energy, prana, are different products and by-products of the second mode of nature, called passion. This mode of passion, however, is the product of the air element, as described before.